But wouldn't the same hold true for humans? Shouldn't their Full Zoan forms be "imperfect"? Or, perhaps the Devil Fruits were never intended for non-human consumption, and that's why Chopper's Zoan cannot give him a "perfect" Zoan form? If you'd like to see the faulty logic behind this assertion, click the spoiler below.
! I've believed for the longest time now that Devil Fruits are people. Shout it with me now, in your best Charlton Heston imitation: DEVIL FRUITS ARE PEOPLE!
! Well, their souls, to be more precise. I'd wondered about this possibility since Robin's flashback and Clover's mentioning of the Lost Kingdom. Even if such a huge kingdom had been destroyed, it would have required wiping out every last one of its inhabitants to create a blank page in history like the Void Century. I wondered where all of those people could have gone to. It would be difficult to cover up mass murder on such a scale so completely.
! Unless the bodies were accounted for as something other than bodies…
! Switching gears for a moment:
! Remember when Lola's crew remarked on the number of shadows Luffy was able to absorb? Moria was able to use shadows as imitation souls to animate his zombies. Brook's DF let his soul come back and reanimate his own body. Between the shadow absorption increasing a person's strength and DFs that allow for limited "soul control", as well as the limitations most people had (the Risky Bros. stated that the most people typically could withstand was two or three) it got me to thinking.
! Why can most people only hold two or three? Is it painful otherwise?
! From this, I interpreted that a shadow contains a fractional amount of a complete soul's energy. Thus, a shadow could reanimate a corpse but not provide it a will or personality of its own- remember, the zombies only retained quirks of the personality they inherited from the shadow's original owner, and usually only for a brief time before coming completely under Moria's will. Lola, Brook, Luffy, Sanji, and Zoro were all good examples of this. However, Zoro and Sanji's personalities seemed to fade much quicker from their respective zombies than Luffy's, Lola's, or Brook's. Why? I thought that in Brook's case, it might be because of the nature of his Devil Fruit- since his own soul was able to reanimate his corpse with his original personality and will, perhaps his shadow retained some of this ability as well? But, this didn't make a good explanation for Luffy or Lola. Unless it was either strength of personality, or the nature of Devil Fruits in general. For the latter idea to hold true, it would require that Lola also have a DF; something which was neither confirmed nor disproven in the story.
! To continue the train of thought, I decided to work under the assumption that Lola did indeed have a DF. So, what then? How did having a DF allow a person's personality to persist in their shadow? Perhaps it had to do with the amount of energy transferred into the corpses? Did DF users have souls with more energy than non-DF users?
! It was during that time that a massive train wreck occurred in the switching station of my mind. The question about whether the member-nations of the WG had turned the people of the Lost Kingdom into fruits collided head on with the question about why Lola, Luffy, and Brook had shadows with persistent personality traits.
! Devil Fruits being the souls of the Lost Kingdom's citizens could explain the complete loss of historical evidence for such a kingdom ever existing. It could also explain why DF users' shadows had more energy than non-DF users. Factoring in the limitations of how many shadows a person could absorb, as well as Blueno's statement that eating two DF caused a person to explode, I concluded that:
! 1. Devil Fruits are the souls of the Lost Kingdom's citizens.
! and
! 2. A complete soul has far more energy than a shadow.
On topic, I still think Kaidou branded the three of them with "Kilroy was here", and Luffy recognized it because it reminded him of Usopp.